刺齿贯众

Cyrtomium caryotideum (Wall. ex Hook. et Grev.) Presl, Tent. Pterid. 86. T. 2. f. 26, 1836; Bedd. Ferns S. Ind. 41. pl. 119. 1863; J. Sm. Hist. Fil. 205. 1875; C. Chr. in Amer. Fern Journ. 20 (2): 50. 1930 et U. S. Nat. Herb. 26 (6): 289. 1931; Tagawa in Acta Phytotax. Geobot. 3 (2): 64 f. 3 (1-4) 1934; Ching in Bull. Chin. Bot. Soc. 2 (2): 99. 1936; Tard.-Blot. et C. Chr. Fl. Indo-Chine 7 (2): 377. 1941; H. Ito, Fil. Jap. Ill. t. 321. 1944;傅书遐, 中国主要植物图说·蕨类植物门194.1957; Tagawa, Col. Ill. Jap. Pterid. 85. 196. pl. 28. f. 165. 1959; Kurata in Sci. Rep. Yokosuka City Mus. 8: 26. 43. 1963; Shing in Acta Phytotax. Sinica Addit. I: 41. 1965; Ic. Corm. Sin. 1: 228. f. 456. 1972; Fl. Tsinling 2: 152. t. 37. f. 7-9. 1974; W. C. Shieh in H. L. Li et al. Fl. Taiwan. 1: 367. 1975; Kurata et Nakaike, Ill. Pterid. Jap. 1: 408. 1979; Dhir, Ferns N. -W. Himal. 78. 1980; Nakaike, N. Fl. Jap. Pterid. 352. 1982; Dixit, Census Ind. Pterid. 140. 1984; Iwatsuki in Ohba et B. Malla, Himal. Pl. 1: 299. 1988; Vasudeva, Bir et Kachroo in Indian Fern J. 7: 79. 1990; S. F. Wu in Invest. Stud. Nat. 12: 33. 1992; Shing in Fl. Jiangxi 1: 146. 1993; Nakaike et Malik in Cryptog. Fl. Pakistan 2: 377. 1993; X. Cheng in W. T. Wang, Vasc. Pl. Hengduan Mts. 1: 146. 1993.——Aspidium cayrotedium Wall. ex Hook. et Grev. Ic. Fil. 1: t. 69. 1828; Hook. Sp. Fil. 4: 40. 1862 et Gard. Ferns pl. 13. 1862.——Aspidium. falcatum Sw. var. caryotideum (Wall.) Hook. et Baker, Syn. Fil. 257. 1867; Franch. Pl. David. 2: 153. 1888.——Cyrtomium falcatum Preslvar. caryotidum (Wall.) Bedd. Handb. Ferns Brit. Ind. 211. 1883; Baroni et Christ in Nouv. Giron. Bot. Ital. n. ser. 4: 93. 1897; Christ in Bull. Soc. Bot. France 52. Mem. 1: 32. 1905; C. Chr. in Acta Hort. Gothob. 1: 72. 1924.——C. caryotideum Preslvar. attenuatum Moore in Gard. Chron. n. s. 1886: 787. ——Polystichum cary otideum (Wall.) Diels in Engl. u. Prantl. Nat. Pflanzfam. 1 (4): 194. 1899. ——Cyrtomium falcatum Presl subvar. hastosum Christ in Bull. Soc. Bot. France 52. Mem. 1: 32. 1905.——C. caryotideum Presl f. attenuatum (Moore) Ching in Bull. Chin. Bot. Soc. 2 (2): 101. 1936; Shing in Acta Phytotax. Sinica Addit. I: 43. 1965. ——C. caryotideum Presl f. hastosum (Christ) Ching 1. c. 101. 1936; Shing 1. c. 43; X. Cheng in W. T. Wang, Vasc. Pl. Hengduan Mts. 1: 147. 1993.

f. caryotideum

植株高30-60厘米。根茎直立，密被披针形黑棕色鳞片。叶簇生，叶柄长16-32厘米，基部直径2-3厘米，禾秆色，腹面有浅纵沟，下部密生卵形及披针形黑棕色或棕色中间为黑棕色的鳞片，鳞片边缘有睫毛状齿，向上部渐秃净；叶片矩圆形或矩圆披针形，长25-48厘米，宽12-18厘米，先端钝，基部不变狭或略变狭，奇数一回羽状；侧生羽片3-7对，互生，斜向上，柄极短，卵状披针形，常向上弯成镰状，中部的长9-14厘米，宽2.5-3.5厘米，先端渐尖常成尾状，基部宽楔形或圆楔形上侧有长而尖的三角形耳状凸起，边缘有开张的小尖齿；具羽状脉，小脉联结成多行网眼，腹面不明显，背面微凸；顶生羽片卵形或菱状卵形，2叉或3叉状，长10-16厘米，宽8-11厘米。叶为坚纸质，腹面光滑，背面疏生披针形棕色小鳞片；叶轴腹面有浅纵沟，疏生线形棕色鳞片。孢子囊群遍布羽片背面；囊群盖圆形，盾状，边缘有齿。

产陕西南部、甘肃南部、江西（井冈山）、台湾（阿里山、台中、花莲）、湖北西部、湖南、广东北部、四川（东部及西南）、贵州、云南、西藏（墨脱）。生林下，海拔600-2500米。也分布于菲律宾、日本、越南、尼泊尔、不丹、印度、巴基斯坦。

f. grossedentatum Ching ex Shing in Acta Phytotax. Sinica Addit. I: 43. 1965; Ching et al. in C. Y. Wu, Fl. Xizang 1: 202. 1983; X. Cheng in W. T. Wang, Vasc. Pl. Hengduan Mts. 1: 147. 1993.

羽片边缘分裂或成粗齿状。

产四川（城口、峨眉山、芦山）、云南（开远、中甸、贡山）、西藏（波密、吉隆、察隅）。生林下岩石上，海拔1200-2700米。模式标本采自西藏波密。