友水龙骨

Polypodiodes amoena (Wall. ex Mett.) Ching in Acta Phytotax. Sin. 16 (4): 27. 1978; Ching et al. in C. Y. Wu, Fl. Xizang. 1: 293. 1983; C. F. Zhang et S. Y. Zhang Fl. Zhejiang. 1: 327. f. 1-347. 1993; S. X. Xu in J. F. Cheng G. F. Chu, Fl. Jiangxi 1: 305. f. 312. 1993. ——Polypodium amoenum Wall., Cat. no. 290. 1828 nom. nud.;Mett., Farngatt. Polypod. 80. 131. 1857; Hook., Sp. Fil. 5: 24. 1863; Syn. Fil. 341. 1868; C. B. Clarke in Trans. Linn. Soc. London, ser. 2. 1: 550. 1880; Diels in Engl. et Prantl, Nat. Pfl. 1: 311. 1899; Christ in Bull. Acad. Geogr. Bot. 1: 203. 1902; C. Chr., Ind. Fil. 508. 1906; Acta Horti Gotob. 1: 99. 1924; Contr. U. S. Nat. Herb. 26: 316. 1931; Wu in Bull. Dept. Biol. Sunyatsen Univ. 3: 264. Pl. 123. 1932; Ching in Contr. Inst. Bot. Nat. Acad. Peiping 2 (3): 43. 1933; Ic. Fil. Sin. 2: ad pl. 99. 1934; Tagawa in Acta Phytotax. Geobot. 8: 234. 1939; Tardieu et Chr. in Fl. Indo-Chine 7 (2): 536. 1941; B. Z. Ding et al., Fl. Henan. 1: 118. 1981; Fl. Shanxi. 1: 124. Pl. 72. F. 1-2. 1992: Tagawa et K. Iwats., Fl. Thailand 3 (4): 569. 1989; De Vol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 202. 1975; Shieh De Vol et C. M. Kuo, Fl. Taiwan 2nd ed. 1: 507. 1994; Shing in W. T. Wang, Vasc. Pl. Hengduan Mt. 1: 160. 1993. ——Goniophlebium amoenum (Wall. ex Mett.) Bedd., Ferns Brit. Ind. 1: ad pl. 5. 1866 et Handb. 317. f. 170. 1883; Rodl-Linder in Blumea 34 (2): 387. 1990. ——Polypodium arisanense Hayata, Ic. Pl, Form. 4: 243. f. 170. 1914. ——Polypodium valdealatum Christ in Bull. Herb. Boiss. 7: 4. 1899. ——Polypodium amoenum Wall. ex Mett. var. latedetoideum Christ in Bull. Acad. Geogr. Bot. 1907: 1421. 1907. ——Polypodium aureum D. Don, Prod. Fl. Nepal. 3. 1825 auct., non Linn., 1753. ——Polypodium bonatianum Brause in Hedwigia 54: 207. Pl. 3. f. 1. 1914. ——Polypodium yunnanense Francht in Bull. Soc. Bot. France 32: 29. 1885; Christ in Bull. Soc. Bot. France 52. Mem. 1: 13. 1905. ——Goniophlebium yunnanense (Franch.) Bedd., Handb. Suppl. 88. 1892; Ching in Contr. Inst. Bot. Nat. Acad. Peiping 2 (3): 46. 1933.

Polypodiodes amoena (Wall. ex Mett.) Ching var. amoena

附生植物。根状茎横走，直径约5-7毫米，密被鳞片；鳞片披针形，暗棕色，基部阔，盾状着生，上部渐尖，边缘有细齿。叶远生；叶柄长约30-40厘米，禾秆色，直径约3-4毫米，光滑无毛；叶片卵状披针形，长约40-50厘米，宽约20-25厘米，羽状深裂，基部略收缩，顶端羽裂渐尖；裂片约20-25对，披针形，长约10-13厘米，宽约1.5-2厘米，顶端渐尖，边缘有锯齿，基部1-2对裂片向后反折。叶脉极明显，网状，在叶轴两侧各具1行狭长网眼，在裂片中脉两侧各具1-2行网眼，内行网眼具内藏小脉，分离的小脉顶端具水囊，几达裂片边缘。叶厚纸质，干后黄绿色，两面无毛，背面叶轴及裂片中脉具有较多的披针形、褐色鳞片。孢子囊群圆形，在裂片中脉两侧各1行，着生于内藏小脉顶端，位于中脉与边缘之间，无盖。

产云南、西藏、四川、贵州、广西、广东、湖南、湖北、江西，浙江、安徽、台湾劝山西。附生于石上或大树干基部，海拔1000-2500米。越南、老挝、泰国、缅甸、印度、尼泊尔、锡金、不丹也有分布。模式标本采自尼泊尔。

Polypodiodes amoena (Wall. ex Mett.) Ching var. duclouxi (Christ) Ching, comb. nov. ——Polypodium duclouxi Christ apudLecomte in Not. Syst. 1: 34. 1909; C. Chr., Ind. Fil. Suppl. 1: 59. 1912; Ching, Ic.Fil. Sin. 2: ad pl. 99. 1934. ——Polypodium amoenum Wall. ex Mett. var. duclouxi (Christ) Ching in Contr. Inst. Bot. Nat. Acad. Peiping 2 (3): 46. 1933; Ic. Corm. Sin. 1: 275. 1972; Shing in W. T. Wang, Vasc. Pl. Hengduan Mt. 1: 160. 1993.

与原变种的区别在于该变种叶柄及叶轴背面常呈紫红色，裂片狭披针形，基部1对裂片不向后反折，叶脉纤细而不甚明显。

产云南和四川。模式标本采自云南。

Polypodiodes amoena (Wall. ex Mett.) Ching var. pilosa (C. B. Clarke) Ching in C. Y. Wu, Fl. Xizang. 1: 294. 1983; C. F. Zhang et S. Y. Zhang, Fl. Zhejiang. 1: 328. 1993. ——Polypodium amoenum Wall. ex Mett. var. pilosum C. B. Clarke in J. Linn. Soc. 24: 417. 1888; Bedd., Handb. Suppl. 89. 1892; Ching in Contr. Inst. Bot. Nat. Acad. Peiping 2 (3): 45. 1933; Ic. Corm. Sin. 1: 275. 1972; Shing in W. T. Wang, Vasc. Pl. Hengduan Mts. 1: 160. 1993. (Type from Khasia). ——Polypodium amoenum Wall. ex Mett. var. pilosum Rosenst. in Fedde Repert. Sp. Nov. 13: 134. 1914. (Type from Guizhou).

与原变种的区别在于该变种叶两面被毛或至少在叶轴及裂片中脉疏被短柔毛。

产云南、西藏、四川、贵州、湖北、浙江。印度北部、尼泊尔、锡金等也有。模式标本采自印度东北部。